Louis R. Joslyn and Marissa Renardy are co-first authors. 

L. R. Joslyn · M. Renardy · C. Weissman · D. E. Kirschner (*) Department of Microbiology and Immunology, University of Michigan Medical School, Ann Arbor, MI, USA e-mail: kirschne@umich.edu 

N. L. Grant · J. L. Flynn Department of Microbiology and Molecular Genetics, University of Pittsburgh School of Medicine, Pittsburgh, PA, USA

J. R. Butler Department of Biology, AdventHealth University, Orlando, FL, USA